Fellcvwebversion.doc

Lloyd Richmond Fell
Academic Qualifications:
Bachelor of Rural Science (University of New England) Master of Rural Science (University of New England) Doctor of Philosophy (University of Melbourne) Thesis on The Neuroendocrine Regulation of Prolactin Graduate Diploma of Social Ecology (University of Western Sydney) Major Project: Stress the Musical
Recent Positions Held:

Senior Research Scientist and Research Leader – NSW Agriculture, and Honorary Fellow in the Centre for Neuroscience and Animal Behaviour University of New England, Armidale, NSW, Australia
Membership of Scientific Societies included:
International Society of Applied Ethology International Society of Comparative Psychology The Scientific and Medical Network The Australasian Society for the Study of Animal Behaviour The Society for Constructivism in the Human Sciences The Personal Construct Psychology (PCP) Interest Group
Research Areas (including over $2 million of research grants awarded):
Stress, behaviour, health and welfare
Cognition and communication
Second-order cybernetics in relation to both the above
Publications:

Approximately 160 published papers, articles or chapters, 60 of these in scientific journals.
The majority prior to 1987 were on aspects of neuroendocrinology or animal husbandry; since
then mostly on animal stress in relation to immunity, health and welfare. Many were on animal
behaviour, in the last decade dealing with aspects of cognition which arose from the biological
research of Maturana and Varela. Since 1991 this has included papers on aspects of human
cognition and behaviour, particularly related to communication and education.

Representative papers are listed below:


Selected Journal Papers (on neuroendocrinology and endocrinology)

Fell L.R. and Mullord M. (1966) Methods of bioassay of oxytocin and vasopressin. N. Z. Med. J. 65: 787.
Fell L.R., Blockey M.A. de B., Beck C., Catt K.J., Brown J.M., Cumming I.A. and Goding J.R. (1971) Prolactin in the
dairy cow during suckling and machine milking. J. Reprod. Fertil. 24: 144.

Fell L.R., Beck C, Brown J.M., Cumming I.A. and Goding J.R. (1972) Radioimmunoassay of ovine prolactin. The
secretion of prolactin as affected by milking, oestradiol administration and onset of parturition. J. Reprod. Fertil. 28:
133.

Fell L.R., Beck C., Brown J.M., Catt K.J., Cumming I.A. and Goding J.R (1972) Solid-phase radioimmunoassay of
ovine prolactin in antibody-coated tubes. Prolactin secretion during estradiol treatment, at parturition and during milking.
Endocrinol. 91: 1329.
Goding J.R., Buckmaster J.M., Cerini J.C., Cerini M.E.D., Chamley W.A., Cumming I.A., Fell L.R., Findlay J.K. and
Jonas H. (1973) Gonadotrophins in the ovine oestrus cycle. J. Reprod. Fertil. Suppl. 18: 31.
Cummins L., Blockey M.A. de B., Fell L.R. and Goding J.R. (1973) Plasma prolactin release about oestrus and after
suckling in the beef cow. J. Reprod. Fertil. 32: 319.
Fell L.R., Perry G.S., Chandler N.J. and Goding J.R. (1973) Effect of the interval between milkings on prolactin release
and milk production in the cow. J. Reprod. Fertil. 32: 317.

Fell L.R., Findlay J.K., Cumming I.A. and Goding J.R. (1973) The effect of synthetic TRF on prolactin release in sheep.
Endocrinol. 93: 215.

Fell L.R., Chandler N.J. and Goding J.R. (1974) Effects of inhibition of prolactin secretion by 2-Br-a-ergocryptine at
parturition and during lactation in dairy cows. J. Reprod. Fertil. 36: 480.
Chamley W.A., Fell L.R., Alford F.P. and Goding J.R. (1974) Twenty-four hour secretion profiles of prolactin and
growth hormone. J. Endocrinol. 61: 165.
Fulkerson W.J., Fell L.R. and McDowell G.H. (1975) Artificial induction of lactation in ewes: the role of prolactin.
Aust. J. Biol. Sci. 28: 525.
Fulkerson W.J., Hooley R.D., McDowell G.H. and Fell L.R. (1976) Progesterone and induction of lactation in ewes. J.
Reprod. Fertil. 46: 512.
Fulkerson W.J., Hooley R.D., Fell L.R. and McDowell G.H. (1976) Artificial induction of lactation in ewes: the
involvement of progesterone and prolactin in lactogenesis. Aust. J. Biol. Sci. 29: 357.
Fulkerson W.J., McDowell G.H., Hooley R.D. and Fell L.R. (1977) Artificial induction of lactation in ewes: the use of
prostaglandin. Aust. J. Biol. Sci. 30: 573.
Hooley R.D., Fell L.R. and Findlay J.K. (1977) The effect of 2-bromo-α-ergocryptine and
2-chloro-α-methylergoline-8β-acetonitrile (lergotrile mesylate) on the prolactin secretion of the ewe. Acta. Endocrinol.
86: 722.



Major Scientific Review Articles (on animal husbandry)

Fell L.R. (1964) Machine milking and mastitis - a review. Dairy Sci. Abst. 26: 551.
Wheeler S.M. and Fell L.R. (1983) Fluorides in cattle nutrition. Nutr. Abstr. Review 53: 741.
Selected Journal Papers (on the biology of stress)

Shutt D.A. and Fell L.R. (1985) Comparison of total and free cortisol in bovine serum and milk or colostrum. J. Dairy
Sci. 68: 1832.

Fell L.R., Shutt D.A. and Bentley C.J. (1985) Development of a salivary cortisol method for detecting changes in
plasma free cortisol arising from acute stress in sheep. Aust. Vet. J. 62: 403.

Fell L.R., Wells R. and Shutt D.A. (1986) Stress in calves castrated surgically or by the application of rubber rings.
Aust. Vet. J. 63: 16.

Fell, L.R. and Shutt, D.A. (1986). Adrenocortical response of calves to transport stress as measured by salivary
cortisol. Can. J. Anim. Sci. 66: 637-641.
Shutt, D.A., Fell, L.R., Connell, R., Bell, A.K., Wallace, C.A. and Smith, A.I. (1987). Stress-induced changes in
plasma concentrations of immunoreactive β-endorphin and cortisol in response to routine surgical procedures in lambs.
Aust. J. Biol. Sci. 40: 97-103.
Shutt, D.A. Fell, L.R., Connell, R. and Bell, A.K. (1988). Stress responses in lambs docked and castrated surgically or
by the application of rubber rings. Aust. Vet. J. 65: 5-7.
Shutt, D.A., Smith, A.I., Wallace, C.A., Connell, R. and Fell, L.R. (1988). Effect of myiasis on plasma immunoreactive
β-endorphin, ACTH, and cortisol in the sheep. Aust. J. Biol. Sci. 41: 297-301.

Fell, L.R. and Shutt, D.A. (1989). Behavioural and hormonal responses to acute surgical stress in sheep. Appl. Anim.
Behav. Sci. 22: 283-294.
Shutt D.A., Connell R. and Fell L.R. (1989) Effects of ovine corticotrophin-releasing factor and vasopressin on plasma
β-endorphin, cortisol and behaviour after minor surgery in sheep. Life Sci. 45: 257-262.
Adams D.B., Lynch J.J., Anderson B.H., Fell L.R., Hinch G.N. and Munro R.K. (1990) The intensity of resistance by
mature Merino ewes against Haemonchus contortus and Trichostrongylus colubriformis in single-species and
combined-species infection. Aust. Vet J. 67 : 443-445.
Fell L.R., Lynch J.J., Adams D.B., Hinch G.N., Munro R.K. and Davies H.I. (1991) Behavioural and physiological
effects in sheep of a chronic stressor and a parasite challenge. Aust. J. Agric. Res. 42 : 1335-1346.
Chapman, R.E., Fell, L.R. and Shutt, D.A. (1994) A comparison of stress in surgically and non-surgically mulesed
sheep. Aust. Vet J. 71 : 243-247.
Adams, D.B. and Fell, L.R. (1997) The effect of infection with the abomasal nematode, Haemonchus contortus, on the
avoidance behaviour of sheep in a motivational-choice test. Int. J. Parasitol. 27 : 665-673.



Selected Conference Papers (on neuroendocrinology and endocrinology)
Fell L.R., Blockey M.A. de B., Beck C., Catt K.J., Brown J.M., Cumming I.A. and Goding J.R. (1970) The control of
milk secretion: levels of prolactin during suckling and milking in dairy cows. Proc. xviii Int. Dairy Congr. 1E: 590.

Fell L.R., Blockey M.A. de B., Beck C., Brown J.M., Cumming I.A., Winfield C.G. and Goding J.R. (1970) Circulating
levels of prolactin in cows and sheep. Proc. 4th Asia Oceania Congr. Endocrinol., p.91.

Fell L.R., Mullord M. and Whittlestone W.G. (1970a) The control of milk ejection: levels of oxytocin and vasopressin
during suckling and milking in the dairy cow. Proc. xviii Int. Dairy Congr. 1E: 591.

Fell L.R., Mullord M. and Whittlestone W.G. (1970b) Circulating levels of oxytocin and vasopressin in the cow. Proc.
4th Asia Oceania Congr. Endocrinol., p.92.
Goding J.R., Buckmaster J.M., Cerini J.C., Cerini M.E.D., Chamley W.A., Cumming I.A., Fell L.R., Findlay J.K. and
Jonas H. (1972) Control of luteal function. Excerpta Medica Int. Cong. Series 273, p. 927.

McCloghry C.E., Fell L.R. and Drinan J.F. (1980) Oestrone, oestradiol-17β, prolactin, α-lactalbumin and lactose
concentration in the mammary secretion of dairy cows around parturition. Proc. Aust. Soc. Reprod. Biol. 12: 89.

Fell L.R. and McCloghry C.E. (1982) The effect of synthetic ACTH administered during late gestation on ovine
placental lactogen and milk secretion. Proc. Aust. Soc. Reprod. Biol. 14: 12.
Fell L.R. and Shutt D.A. (1986) Use of salivary cortisol as an indicator of stress due to management practices in sheep
and calves. Proc. Aust. Soc. Anim. Prod. 16: 203.
Shutt D.A., Fell L.R., Connell R., Wallace C.A. and Smith A.I. (1986) Changes in plasma cortisol and β-endorphin in
response to routine surgery in lambs. Proc. Endocrinol. Soc. Aust. Abstract 104.

Fell L.R. (1987) Behavioural responses to husbandry stressors in lambs. Proc. Aust. Soc. Anim. Behav., p. 12.
Shutt, D.A. and Fell, L.R. (1988). The role of endorphins in the response to stress in sheep and cattle. Proc. Aust. Soc.
Anim. Prod. 17: 338-341.

Fell, L.R. and Shutt, D.A. (1988). Salivary cortisol and behavioural indicators of stress in sheep. Proc. Aust. Soc.
Anim. Prod. 17: 186-189.

Fell L.R. and Shutt D.A. (1989) Development of an arena test to measure aversion in stressed sheep. Proc. 16th Ann.
Conf. Australasian Soc. Anim. Behav., p.7.

Shutt D.A., Fell L.R., Adams D.B. and Lynch J.J. (1990) Effects of Haemonchus contortus in pen-fed versus paddock
sheep as determined by plasma cortisol and insulin-like growth factor-1. Proc. Aust. Soc. Anim. Prod. 18 : 360-363.

Fell L.R., Lynch J.J., Adams D.B., Hinch G.N. and Munro R.K. (1990) The behavioural response of immune sheep to a
challenge infection with an internal parasite. Proc. 17th Ann. Conf. Australasian Soc. Anim. Behav., p. 10.
Barnett J.P, Fell L.R., Lynch J.J., Adams D.B., Hinch G.N., Munro R.K. and Gates G.R. (1990) Effect of opiate
antagonist and antihistamine on behaviour of immune ewes infected with Haemonchus contortus. Proc. 17th Ann.
Conf. Australasian Soc. Anim. Behav., p. 11.

Fell L.R., Lynch J.J., Adams D.B., Hinch G.N. and Munro R.K. (1990) Effects of a challenge infection with
Haemonchus contortus on the behaviour of penned versus paddock sheep. Proc. Aust. Soc. Anim. Prod. 18 : 475.

Fell L.R., Lynch J.J., Adams D.B., Hinch G.N., Barnett J.P., Gates G.R. and Bertus F.V. (1990) Behaviour following
challenge infections with Haemonchus contortus in immune and non-immune sheep. Proc. Scientific Meeting, Aust.
Behavioural Immunol. Group, p. 4.
Adams D.B., Fell L.R., Lynch J.J., Barnett J.P., Gates G.R. and Hinch G.N. (1990) The time-course of behavioural
effects of immune responses to challenge infection with Haemonchus contortus in sheep. Proc. 1st Scientific Meeting,
Aust. Behavioural Immunol. Group, p. 6.
Barnett J.P., Adams D.B., Lynch J.J., Fell L.R., Hinch G.N., Munro R.K. and Gates G.R. (1990) Evidence for a link
between endogenous opiate and immune systems in sheep infected with Haemonchus contortus. Proc. 1st Scientific
Meeting, Aust. Behavioural Immunol. Group, p. 5.
Barnett J.P., Gates G.R., Lynch J.J., Adams D.B., Fell L.R., Hinch G.N., Davies H.I. and Gregson R.A.M. (1991)
Interpretation of motivational mechanisms of sheep in an arena test. Proc. 18th Ann. Conf. Australasian Soc. Anim.
Behav. p. 8.
Behrendt R., Fell L.R., Newman R., Rigby R.D.G., Hoskinson R.M. and Wynn P.C. (1991) Active immunisation
against ACTH (1-24) modifies the behaviour of merino sheep. Proc. Scientific Meeting Aust. Endocrin. Soc. Abstract
143, p. 111.
Degabriele R. and Fell L.R. (1992) Interleukin-1 production and behavioural adaptation. Proc. 2nd Scientific Meeting,
Aust. Behavioural Immunol. Group, p. 12.

Fell, L.R. and Clarke, M.R. (1993) Behaviour of lot-fed cattle. Recent Advances in Animal Nutrition in Australia 1993.
Dept. Biochem., Microbiol. and Nutrition, Univ. of New England, Armidale, NSW, Australia. pp 107-116.
Watson, D.L. and Fell, L.R. (1993) Stress and immune competence in feedlot cattle. Recent Advances in Animal
Nutrition in Australia 1993. Dept. Biochem., Microbiol. and Nutrition, Univ. of New England, Armidale, NSW,
Australia. pp 130-136.



Selected Journal and Conference Papers (on cognition and communication)

Fell, L.R. and Russell, D.B. (1991) The practice of science: the research-development relationship with particular
reference to agriculture. Proc. Gadamer Action and Reason Conf. University of Sydney, pp. 25-30.
Fell L.R. (1991) Sheep behaviour: does it matter what sheep think? Rural Science Annual, University of New England,
Armidale, pp. 45-47.
Ison, R.L., Russell, D.B. and Fell, L.R. (1993) Knowledge - You can't give it away. Proc. Universities as
Interpretive Communities Conf., University of Sydney, pp. 169-174.
Fell, L.R. and Russell, D.B. (1994) Towards a biological explanation of human understanding. J. Cybernetics and
Human Knowing 2 : (4) 3-15.

Book Chapters (on stress/cognition and Maturana’s biological research)
Fell L.R. and Shutt D.A. (1989) The concept of stress in farm animals - a review. In Rose M. and Large A. (Eds)
Animal Ethics Committees - The Role of Veterinarians, p. 66.
Gates G.R., Fell L.R., Lynch J.J., Adams D.B., Barnett J.P., Hinch G.N., Munro R.K. and Davies H.I. (1992) The link
between immune responses and behaviour in sheep. Chapter 3 in Husband A.J. (Ed) Behaviour and Immunity, CRC
Press, Boca Raton, pp. 23-41.

Fell, L.R. and Russell, D.B. (1994) Biology's Room With a View. In Fell, L.R., Russell, D.B. and Stewart, A.H.
(Eds) Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the visit to Australia
in August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University of Western
Sydney, Richmond, NSW, Australia.
Fell, L.R. and Russell, D.B. (1994) An Introduction to "Maturana's" Biology. In Fell, L.R., Russell, D.B. and
Stewart, A.H. (Eds) Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the
visit to Australia in August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University
of Western Sydney, Richmond, NSW, Australia.
Fell, L.R. and Russell, D.B. (1994) Living Systems - Autonomous Unities. In Fell, L.R., Russell, D.B. and Stewart,
A.H. (Eds) Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the visit to
Australia in August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University of
Western Sydney, Richmond, NSW, Australia.
Fell, L.R. and Russell, D.B. (1994) Non-Traditional R & D. In Fell, L.R., Russell, D.B. and Stewart, A.H. (Eds)
Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the visit to Australia in
August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University of Western Sydney,
Richmond, NSW, Australia.
Fell, L.R. (1994) Stress, Epistemology and Feedlot Cattle. In Fell, L.R., Russell, D.B. and Stewart, A.H. (Eds)
Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the visit to Australia in
August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University of Western Sydney,
Richmond, NSW, Australia.
Russell, D.B., Dimitrov, V. and Fell, L.R. (1994) Ancient Wisdom and Contemporary Ecological Problems. In Fell,
L.R., Russell, D.B. and Stewart, A.H. (Eds) Seized by Agreement, Swamped by Understanding - A collection of
papers to celebrate the visit to Australia in August 1994 by Humberto Maturana. ISBN 0 646 20084 4 -
Hawkesbury Printing - University of Western Sydney, Richmond, NSW, Australia.
Fell, L.R. and Russell, D.B. (1994) The Dance of Understanding. In Fell, L.R., Russell, D.B. and Stewart, A.H.
(Eds) Seized by Agreement, Swamped by Understanding - A collection of papers to celebrate the visit to Australia
in August 1994 by Humberto Maturana. ISBN 0 646 20084 4 - Hawkesbury Printing - University of Western
Sydney, Richmond, NSW, Australia.
Fell, L.R. (1996) The Struggle to Integrate. In Ryan, J.S. (Ed.) Rural Science: Philosophy and Application. School
of Rural Science, University of New England, Armidale, NSW, Australia, p 464-467.

Fell, L.R. and Russell, D.B. (in press - 1998) The Human Quest for Understanding and Agreement. Chapter 3 in
Ison, R. and Russell, D.B. (Eds) Breaking With Tradition – A New Approach to Agricultural Extension..
Cambridge University Press, London, UK.




Recent Student Supervision, Examining and Teaching Activities:
Robert Degabriele - PhD - University of Western Sydney, Macarthur - Stress and Immunity
Stephen Wilson - PhD - University of Wollongong - Indicators of Animal Welfare
Honours students – Centre for Neuroscience and Animal Behaviour, University of New
England.
I have supervised numerous Masters and Honours students from the University of Sydney,
University of NSW and University of Western Sydney, Hawkesbury over the last 10 years and
examined about a dozen postgraduate theses including one PhD thesis from Ronald Proudford
at the University of NSW on Public Health Administration.
I have served as an invited lecturer on many occasions at the University of Sydney, University
of NSW, University of Western Sydney and University of New England, mainly on stress
physiology and behaviour and, more recently, on second-order cybernetics

Other Professional Activities:
External reviewer of the Department various University Departments.
External reviewer of numerous research grant applications in Australia and New Zealand.
External referee for numerous scientific journal papers in Australia and New Zealand.
Committee member of several scientific societies at various times.
Research team leader, Conference organiser, occasional press, TV, radio appearances.
Maintaining the only Australian web site on autopoiesis and the international network
associated with this.
Conducting workshops on the application of Maturana’s biological research and second-order
cybernetics to improving the quality of life of individuals and societies.
Songwriting and performing, particularly in connection with workshops on cognition and
communication.

Source: http://www.pnc.com.au/~lfell/CVweb.pdf